Plant Micronutrient Functions
Nutrient Deficiency Symptoms In Plants
A primary
function of boron is related to cell wall formation, so
boron-deficient plants may be stunted. Sugar transport in
plants, flower retention and pollen formation and germination
also are affected by boron. Seed and grain production are
reduced with low boron supply. Boron-deficiency symptoms first
appear at the growing points. This results in a stunted
appearance (rosetting), barren ears due to poor pollination,
hollow stems and fruit (hollow heart) and brittle, discolored
leaves and loss of fruiting bodies.
Boron
deficiencies are found mainly in acid, sandy soils in regions of
high rainfall, and those with low soil organic matter. Borate
ions are mobile in soil and can be leached from the root zone.
Boron deficiencies are more pronounced during drought periods
when root activity is restricted.
Copper
Copper is
necessary for carbohydrate and nitrogen metabolism and,
inadequate copper results in stunting of plants. Copper also is
required for lignin synthesis which is needed for cell wall
strength and prevention of wilting. Deficiency symptoms of
copper are dieback of stems and twigs, yellowing of leaves,
stunted growth and pale green leaves that wither easily.
Copper
deficiencies are mainly reported on sandy soils which are low in
organic matter. Copper uptake decreases as soil pH increases.
Increased phosphorus and iron availability in soils decreases
copper uptake by plants.
Iron
Iron is
involved in the production of chlorophyll, and iron chlorosis is
easily recognized on iron-sensitive crops growing on calcareous
soils. Iron also is a component of many enzymes associated with
energy transfer, nitrogen reduction and fixation, and lignin
formation. Iron is associated with sulfur in plants to form
compounds that catalyze other reactions. Iron deficiencies are
mainly manifested by yellow leaves due to low levels of
chlorophyll. Leaf yellowing first appears on the younger upper
leaves in interveinal tissues. Severe iron deficiencies cause
leaves to turn completely yellow or almost white, and then brown
as leaves die.
Iron
deficiencies are found mainly on high pH soils, although some
acid, sandy soils low in organic matter also may be
iron-deficient. Cool, wet weather enhances iron deficiencies,
especially on soils with marginal levels of available iron.
Poorly aerated or compacted soils also reduce iron uptake by
plants. Uptake of iron decreases with increased soil pH, and is
adversely affected by high levels of available phosphorus,
manganese and zinc in soils.
Manganese
Manganese is
necessary in photosynthesis, nitrogen metabolism and to form
other compounds required for plant metabolism. Interveinal
chlorosis is a characteristic manganese-deficiency symptom. In
very severe manganese cases, brown necrotic spots appear on
leaves, resulting in premature leaf drop. Delayed maturity is
another deficiency symptom in some species. White/gray spots on
leaves of some cereal crops is a sign of manganese deficienc.
Manganese
deficiencies mainly occur on organic soils, high-pH soils, sandy
soils low in organic matter, and on over-limed soils. Soil
manganese may be less available in dry, well-aerated soils, but
can become more available under wet soil conditions when
manganese is reduced to the plant-available form. Conversely,
manganese toxicity can result in some acidic, high-manganese
soils. Uptake of manganese decreases with increased soil pH and
is adversely affected by high levels of available iron in soils.
Molybdenum
Molybdenum is
involved in enzyme systems relating to nitrogen fixation by
bacteria growing symbiotically with legumes. Nitrogen
metabolism, protein synthesis and sulfur metabolism are also
affected by molybdenum. Molybdenum has a significant effect on
pollen formation, so fruit and grain formation are affected in
molybdenum-deficient plants. Because molybdenum requirements are
so low, most plant species do not exhibit molybdenum-deficiency
symptoms. These deficiency symptoms in legumes are mainly
exhibited as nitrogen-deficiency symptoms because of the primary
role of molybdenum in nitrogen fixation. Unlike the other
micronutrients, molybdenum-deficiency symptoms are not confined
mainly to the youngest leaves because molybdenum is mobile in
plants. The characteristic molybdenum deficiency symptom in some
vegetable crops is irregular leaf blade formation known as
whiptail, but interveinal mottling and marginal chlorosis of
older leaves also have been observed.
Molybdenum
deficiencies are found mainly on acid, sandy soils in humid
regions. Molybdenum uptake by plants increases with increased
soil pH, which is opposite that of the other micronutrients.
Molybdenum deficiencies in legumes may be corrected by liming
acid soils rather than by molybdenum applications. However, seed
treatment with molybdenum sources may be more economical than
liming in some areas.
Zinc
Zinc is an
essential component of various enzyme systems for energy
production, protein synthesis, and growth regulation. Zinc
deficient plants also exhibit delayed maturity. Zinc is not
mobile in plants so zinc-deficiency symptoms occur mainly in new
growth. Poor mobility in plants suggests the need for a constant
supply of available zinc for optimum growth. The most visible
zinc deficiency symptoms are short internodes and a decrease in
leaf size. Delayed maturity also is a symptom of zinc-deficient
plants.
Zinc
deficiencies are mainly found on sandy soils low in organic
matter and on organic soils. Zinc deficiencies occur more often
during cold, wet spring weather and are related to reduced root
growth and activity as well as lower microbial activity
decreases zinc release from soil organic matter. Zinc uptake by
plants decreases with increased soil pH. Uptake of zinc also is
adversely affected by high levels of available phosphorus and
iron in soils.
Chloride
Because
chloride is a mobile anion in plants, most of its functions
relate to salt effects (stomatal opening) and electrical charge
balance in physiological functions in plants. Chloride also
indirectly affects plant growth by stomatal regulation of water
loss. Wilting and restricted, highly branched root systems are
the main chloride-deficiency symptoms, which are found mainly in
cereal crops.
Most soils
contain sufficient levels of chloride for adequate plant
nutrition. However, reported chloride deficiencies have been
reported on sandy soils in high rainfall areas or those derived
from low-chloride parent materials. There are few areas of
chloride-deficient so this micronutrient generally is not
considered in fertilizer programs. In addition, chloride is
applied to soils with KCl, the dominant potassium fertilizer.
The role of chloride in decreasing the incidence of various
diseases in small grains is perhaps more important than its
nutritional role from a practical viewpoint.
Plants differ
in their requirements for certain micronutrients. The following
table shows the estimate of the relative response of selected
crops to micronutrients. The ratings of low medium and high are
used to indicate the relative degree of responsiveness.
Table 1. Crop Response to Micronutrients
Crop
|
B
|
Cu
|
Mn
|
Zn
|
Alfalfa
|
High
|
Med
|
Low
|
Med
|
Wheat
|
Low
|
High
|
High
|
Low
|
Rye
|
Low
|
Low
|
Low
|
Med
|
Clover
|
Med
|
Med
|
Low
|
Med
|
Corn
|
Med
|
Low
|
Low
|
High
|
Oats
|
Low
|
Med
|
High
|
Med
|
Sugar Beet
|
High
|
Low
|
Low
|
Med
|
Grass
|
Low
|
Low
|
Low
|
Med
|
|